Update: I replaced MUTAGEN, in all caps, to mutagen, in all lowers, and that did the trick. Nonetheless I found an error with “MUT_IV” later, but replacing that with “mutagen_iv” was good enough.
Nocts’ Cataclysm++ mod too showed the “MUT_IV” error in tools.json, as well as the mod “Mutation changes” in the file mutations.json (this one also had MUTAGEN in all caps in the same file).